中国科学院院士、教授、博导

E-mail: szshen@nju.edu.cn

地层古生物工作是“穷人”从事的贵族职业，是所有地质学科的基础，地球科学人“放眼世界，胸怀全球”。

一、个人概况

  1961年生，浙江湖州人，1981年毕业于浙江煤炭工业学校，1981-1983曾在浙江省长广煤矿公司查扉矿工作，1986年在中国矿业大学获得硕士学位，1989年在该校获得博士学位。1996-2000分别在日本新泻大学和澳大利亚Deakin大学从事博士后研究，2000年底由中科院“百人计划”从澳大利亚引进到南京地质古生物所工作。曾获国家杰出青年基金(2002)，主持基金委优秀创新群体9年（2004-2012)；任科技部973（2006-2010）、基金委重大（2013-2017）、青藏高原第二次科考计划专题（2019-2023）等项目首席科学家；《现代古生物学和地层学》国家重点实验室主任（2006-2015）。现任《国际地层委员会》副主席和《Palaeogeography, Palaeoclimatology, Palaeoecology》共同主编。在二叠纪地层学、二叠纪末生物大灭绝与环境变化、腕足动物和牙形类古生物学等方面取得了系统性和创新性成果。2015年当选中国科学院院士。

业已发表论著350余篇，出版专著或主编专集25册。野外工作足迹涉及50多个国家。由于在二叠纪地层学方面的突出贡献，2019年被授予国际地层委员会最高金奖(ICS Medal)。还曾获国家自然科学二等奖和省科技进步一等奖各1项（均排名第二）、国家六部委颁发的优秀回国人员成就奖、江苏省高层次人才突出贡献奖、李四光地质科学奖等、江苏省先进工作者等。

二、研究领域

主要从事二叠纪腕足动物、牙形类化石系统古生物学、生物大灭绝及其以后的复苏、定量生物古地理学、生物多样性和古环境学等方面的研究。在系统古生物研究方面，他深入研究了十多个国家的腕足类和牙形类动物群。带领团队建立石炭纪-三叠纪全球腕足动物数据库，创建全球生物古地理和多样性演变模式，使其成为判断该时段地体古地理位置和古海道开闭时间以及海洋底栖生态系统演变依据。

是6个二叠纪“金钉子”研究的主要完成人之一。领衔的课题组与国外实验室合作完成的多个重要界线的高精度年龄等被纳入最新的《国际年代地层表，2022》。1994年第一次赴西藏希夏邦马峰地区进行野外工作以后对喜马拉雅地区二叠纪地质产生浓厚兴趣．先后5次进入西藏进行野外地质工作，分别对藏南地区的色龙剖面、土隆剖面和珠穆朗玛峰地区的多个剖面进行了详细的地层和古生物研究，发表了一系列论著，建立了喜马拉雅地区的高分辨率的二叠系－三叠系地层和地球化学剖面，基本阐明了南方高纬度地区的二叠纪末生物大灭绝模式。

在二叠纪生物大灭绝研究中，与团队成员和国际同行一起共同开展高精度综合年代地层框架、生物多样性定量分析和多种地化指标等研究，论证二叠纪末海陆生物大灭绝的同时性和瞬时性，指出特提斯洋中酸性岩浆弧火山喷发和西伯利亚大规模岩浆活动造成地表环境的巨变是导致该生物大灭绝的主因。相关成果在Science、Science Advances、PNAS、Geology、EPSL等刊物发表后,引起国内外同行较大关注。2011年发表在Science上的成果入选《2012年度中国科学十大进展》，相关内容被编入美国大学教材等。2020年与大数据团队一起，利用大数据平台、开发新算法等共同创建了古生代3亿多年的高分辨率生物多样性演变曲线，成果发表在Science上，分别被评为2020年度《中国科学十大进展》和《中国科学十大进展新闻》等。

  此外，每年到国外开展野外工作，涉及美国、加拿大、欧洲、南非、中国周边的日本、缅甸、巴基斯坦、伊朗、泰国、蒙古等，在国内，分别在新疆、华南、东北、华北等各地进行二叠纪地层和腕足动物、牙形类化石群的研究，特别对新疆和华北含油气、煤炭资源地层、广西、四川、重庆、贵州、云南等地二叠纪地层和腕足动物化石群进行了详细地研究。对建立详细的大型数据库具有浓厚的兴趣。除了腕足动物化石群以外，他还对牙形类化石、菊石类化石、古植物化石等都有一定的兴趣，现有研究生和博士后分别从事这些方面地研究。

三、获奖及荣誉

由于他在二叠纪地层学方面的突出贡献，2019年被授予国际地层委员会最高金奖(ICS Medal)，目前是亚洲唯一获此殊荣的学者。此外，他还曾获国家自然科学二等奖(2010)和省科技进步一等奖(2008)各1项（均排名第二）、国家六部委颁发的优秀回国人员成就奖(2003)、江苏省高层次人才突出贡献奖(2011)、尹赞勋地层古生物学奖(2013)、李四光地质科学奖(2015)、江苏省先进工作者（2021）等。

四、承担科研项目情况

先后主持和参加了10余项国家自然科学基金委、澳大利亚联邦基金委、科技部和原煤炭部的项目。是2000年度中国科学院《引进海外杰出人才计划》（百人计划）获得者，2002年度获得《国家杰出青年基金》。2003年度国家自然科学基金委员会资助的《优秀创新群体》学术带头，2006年作为首席科学家获得科技部973项目《地史时期海陆生物多样性的演变》。2008年和2015年分别获得国家自然基金委重点国际合作项目资助。2013-2017主持基金委重大项目《古生代重大海洋生物事件》。目前正在主持《青藏高原第二次科考计划》专题、中科院专项B课题等。

五、代表性论著

一）、专著和主编专辑

1.        沈树忠、徐海鹏、袁东勋、王玥、王军、张以春、王小娟、牟林、吴琼. 2020. 中国二叠纪地层及标志化石图集。浙江大学出版社，382 p.

2.        Shen, S.Z. & Rong, J.Y. 2019. Integrative Stratigraphy and Timescale of China. Science China Earth Sciences 62(1):1-348(沈树忠、戎嘉余（主编），2019. 中国综合地层和时间框架。中国科学:地球科学, 49(1):1-352)。(Special Issue)

3.        Lucas, S.G. & Shen, S.Z. (eds.). 2018. The Permian Timescale. Geological Society, London, Special Publications 450: 1-458.

4.        Rong, J.Y., Jin, Y.G., Shen, S.Z. & Zhan, R.B. (eds.). 2017. Phanerozoic Brachiopod Genera of China, Volumes 1-2, 1-1077.

5.        Shen, S.Z., Jin, J.S. & Shi, G.R. 2016. Ecosystem evolution in deep time: Evidence from the rich Paleozoic fossil records of China. Palaeogeography, Palaeoclimatology, Palaeoecology 448: 1-382. (Special Issue)

6.        Shen, S.Z., Shi, G.R. 2013. Late Palaeozoic deep Gondwana and its peripheries: stratigraphy, biological events, palaeoclimate and palaeogeography. Gondwana Research 24(1): 1-242. (Special Issue).

7.        Shen, S.Z., Henderson, C.M. & Somerville, I.D. 2010. Lopingian (Late Permian) stratigraphy of the world, major events and environmental change. Geological Journal 45(2-3):119-349. (Special Issue)

8.        Shen, S.Z. & Zhou Zhong-he (eds.). 2010. Evolution of marine and terrestrial geobiodiversity in the history of the earth. Science in China Series D: Earth Sciences, 53(12):1747-1940(沈树忠、周忠和,主编. 2010.地史时期海陆生物多样性演变。中国科学D:地球科学，40(9):1095-1300). (Special Issue)

9.        Shen, S.Z. & Shi G.R. 2007. Lopingian (Late Permian) Brachiopoda from South China, Part 1: Orthotetida, Orthida and Rhynchonellida. Bulletin of the Tohoku University Museum, no. 6:1-102.

10.    Shen, S.Z., Wang, X.D. & Erwin, D.H. 2007. Contributions to brachiopod paleontology, Permian and Carboniferous stratigraphy and end-Permian mass extinctions-in memory of Prof. Jin Yugan. Palaeoworld 16 (1-3):1-263. (Special Issue).

11.    Shi, G.R., Campi, M. A. & Shen, S.Z., (eds.), 2006. The Permian of East and NE Asia. Journal of Asian Earth Sciences 26:173-436 (Special Issue)

12.    Shen, S.Z., Shi, G.R. & Archbold, N.W. 2003. Lopingian (Late Permian) brachiopods from the Qubuerga Formation at the Qubu section in the Mt. Qomolangma region, southern Tibet (Xizang), China. Palaeontographica Band 268, 49-101.

二）、论文

13.    Zhang, H., Zhang, F.F., Chen, J.B., Erwin, D.H., Syverson, D.D., Ni, P., Rampino, M., Chi, Z., Cai, Y.F., Xiang, L., Li, W.Q., Liu, S.A., Wang, R.C., Wang, X.D., Feng, Z., Li, H.M., Zhang, T., Cai, H.M., Wang Z., Cui, Y., Zhu, X.K., Hou, Z.Q., Wu, F.Y., Xu, Y.G., Planavsky, N., Shen, S.Z.* 2021. Felsic volcanism as a factor driving the end-Permian mass extinction. Science Advances 7:eabh1390, 13p.

14.    Wu, Q., Ramezani, J., Zhang, H.*, Wang, J., Zeng, F.G., Zhang, Y.C., Liu, F., Liu, C., Yang, W., Henderson, C.M., Shen, S.Z.* 2021. High-precision U-Pb age constraints on the Permian floral turnovers, paleoclimate change and tectonics of North China. Geology 49:677-681.

15.    Zhang, M., Qin, H.F. He, K., Hou, Y.F., Zheng, Q.F., Deng, C.L., He, Y. & Shen, S.Z.*, Zhu, R.X. & Pan, Y.X.* 2021. Magnetostratigraphy across the end-Permian mass extinction event from the Meishan sections, southeastern China. Geology 49:1289-1294.

16.    Hu, Z.Y., Li, W.Q.*, Zhang, H.*, Krainer, K., Zheng, Q.F., Xia, Z.G., Hu, W.X. & Shen, S.Z. 2021. Mg isotope evidence for restriction events within the Paleotethys ocean around the Permian-Triassic transition. Earth and Planetary Science Letters 556: 116704.

17.    Wang, W.Q.*, Katchinoff, J.A.R., Garbelli, C., Immenhauser, A., Zheng, Q.F., Zhang, Y.C., Yuan, D.X., Shi, Y.K., Wang, J.Y., Planavsky, N. & Shen, S.Z.* 2021. Revisiting the Permian seawater 87Sr/86Sr record: New perspectives from brachiopod proxy data and stochastic oceanic box models. Earth-Science Reviews 218: 103679.

18.    Wang, C.S., Hazen, R.M., Cheng, Q.M., Stephenson, M.H., Zhou, C.H., Fox, P., Shen, S.Z., Oberhänsli, R., Hou, Z.Q., Ma, X.G., Feng, Z.Q., Fan, J.X., Ma, C., Hu, X.M., Luo, B. & Wang, J.L. 2021. The Deep-time Digital Earth program: data-driven discovery in geosciences. National Science Review 8(9): nwab027.

19.    Xu, H.P., Aung, K.P., Zhang, Y.C., Shi, G.R., Cai, F.L., Zaw, T., Ding, L., Sein, K. & Shen, S.Z.* 2021. A late Cisuralian (early Permian) brachiopod fauna from the Taungnyo Group in the Zwekabin Range, eastern Myanmar and its biostratigraphic, paleobiogeographic, and tectonic implications. Journal of Paleontology 95: 1158-1188.

20.    Zhang, S.H.*, Fan, J.X., Morgan, C.A., Henderson, C.M. & Shen, S.Z.* 2021. Quantifying the middle–late Cambrian trilobite diversity pattern in South China. Palaeogeography, Palaeoclimatology, Palaeoecology 570:110361.

21.    Shen, S.Z.*, Yuan, D.X.*, Henderson, C.M., Wu, Q., Zhang, Y.C., Zhang, H., Mu, L., Ramezani, J., Wang, X.D., Lambert, L.L., Erwin, D.H., Hearst, J.M., Xiang, L., Chen, B., Fan, J.X., Wang, Y., Wang, W.Q., Qi, Y.P., Chen, J., Qie, W.K. & Wang, T.T. 2020. Progress, problems and prospects: An overview of the Guadalupian Series of South China and North America. Earth-Science Reviews 211: 103412.

22.    Fan, J.X., Shen, S.Z.* Erwin, D.H., Sadler, P.M., MacLeod, N., Cheng, Q.M., Hou, X.D., Yang J., Wang, X.D., Wang, Y., Zhang, H., Chen, X., Li, G.X., Zhang, Y.C., Shi, Y.K., Yuan, D.X., Chen, Q., Zhang, L.N., Li, C. & Zhao, Y.Y. 2020. A high-resolution summary of Cambrian to Early Triassic marine invertebrate biodiversity. Science 367:272-277.

23.    Planavsky, N.J., Hood, A., Tarhan, L.G., Shen, S.Z. & Johnson, K. 2020. Store and share ancient rocks. Nature 581:137-139.

24.    Wang, W.Q., Garbelli, C., Zhang, F.F., Zheng, Q.F., Zhang, Y.C., Yuan, D.X., Shi, Y.K., Chen, B. & Shen, S.Z.* 2020. A high-resolution Middle to Late Permian paleotemperature curve reconstructed using oxygen isotopes of well-preserved brachiopod shells. Earth and Planetary Science Letters 540:116245.

25.    Zhang, F.F.*, Shen, S.Z.*, Cui, Y., Lenton, T.M., Dahl, T.W., Zhang, H., Zheng, Q.F., Wang, W.Q., Krainer, K. & Anbar, A.D. 2020. Two distinct episodes of marine anoxia during the Permian-Triassic crisis evidenced by uranium isotopes in marine dolomites. Geochimica et Cosmochimica Acta 287:165-179.

26.    Chernykh, V.V., Chuvashov, B.I., Shen, S.Z.*, Henderson, C.M., Yuan, D.X., Stephenson, M.H. 2020. The Global Stratotype Section and Point (GSSP) for the base-Sakmarian Stage (Cisuralian, Lower Permian). Episodes 43(4):961-979.

27.    Yuan, D.X., Aung Kyi Pyar, Henderson, C.M., Zhang, Y.C., Zaw Than, Cai, F.L., Ding, L., Shen, S.Z.* 2020. First records of Early Permian conodonts from eastern Myanmar and implications of paleobiogeographic links to the Lhasa Block and northwestern Australia. Palaeogeography, Palaeoclimatology, Palaeoecology 549:109363.

28.    Hou, Z.S., Fan, J.X., Henderson, C.M., Yuan, D.X., Shen, B.H., Wu, J., Wang, Y., Zheng, Q.F., Z., Zhang, Y.C., Wu, Q. & Shen, S.Z.* 2020. Dynamic palaeogeographic reconstructions of the Wuchiapingian Stage (Lopingian, Late Permian) for the South China Block. Palaeogeography, Palaeoclimatology, Palaeoecology 546:109667.

29.    Wu, Q., Ramezani, J., Zhang, H., Yuan, D.X., Erwin, D.H., Henderson, C.M., Lambert, L.L., Zhang, Y.C. & Shen, S.Z.* 2020. High-precision U-Pb zircon age constraints on the Guadalupian in West Texas, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 548:109668.

30.    Zhou, W.M., Aung, K.P., Liu, L., Zhang, Y.C., Zaw, T., Wang, J. & Shen, S.Z.* 2020. First record of Cisuralian–Guadalupian plant fossils from the Shan Plateau, eastern Myanmar. Palaeoworld 29:108-116.

31.    Henderson, C.M. & Shen, S.Z. 2020. The Permian Period. In: Gradstein, F.M., Ogg, J.G., Schmitz, M.D. & Ogg, G.M., eds., Geologic Time Scale 2020. Amsterdam, Oxford, Cambridge, Elsevier, p. 875-902.

32.    Garbelli, Claudio*, Shen, S.Z.*, Immenhauser, A., Brand, U., Buhl, D., Wang, W.Q., Zhang, H. & Shi, G.R. 2019. Timing of Early and Middle Permian deglaciation of the southern hemisphere: brachiopod-based 87Sr/86Sr calibration. Earth and Planetary Science letters 516:122-135.

33.    Wang, J.Y., Jacobson, A.D., Zhang, H., Ramezani, J., Sageman, B.B., Hurtgen, M.T., Bowring, S.A., Shen, S.Z. 2019. Coupled δ⁴⁴/⁴⁰Ca, δ⁸⁸/⁸⁶Sr, and ⁸⁷Sr/⁸⁶Sr geochemistry across the end-Permian mass extinction event. Geochimica et Cosmochimica Acta 262: 143-165.

34.    Shen, S.Z.*, Ramezani, J., Chen, J.*, Cao, C.Q., Erwin, D.H., Zhang, H., Xiang, L., Schoepfer, S.D., Henderson, C.M., Zheng, Q.F., Bowring, S.A., Wang, Y., Li, X.H., Wang, X.D., Yuan, D.X., Zhang, Y.C., Mu, L., Wang, J., Wu, Y.S. 2019. A sudden end-Permian mass extinction in South China. GSA Bulletin 131:205-223.

35.    Xu, H.P., Zhang, Y.C., Qiao, F. & Shen, S.Z.* 2019. A new Changhsingian brachiopod fauna from the Xiala Formation at Tsochen in the central Lhasa Block and its paleogeographical implications. Journal of Paleontology 93:876-898.

36.    Wu, H.C., Fang, Q., Wang, X.D., Hinnov, L.A., Qi, Y.P., Shen, S.Z., Yang, T.S., Li, H.Y., Chen, J.T. & Zhang, S.H. 2019. A ~34 m.y. astronomical time scale for the uppermost Mississippian through Pennsylvanian of the Carboniferous System of the Paleo-Tethyan Realm. Geology 47:83-86.

37.    Shen, S.Z., Zhang, H., Zhang, Y.C., Yuan, D.X., Chen, B., He, W.H., Mu, L., Lin, W., Wang, W.Q., Chen, J., Wu, Q., Cao, C.Q., Wang, Y. & Wang, X.D. 2019. Permian integrative stratigraphy and timescale of China. Science in China Earth Sciences 62(1):154-188. （沈树忠，张华，张以春，袁东勋，陈波，何卫红，牟林，林巍，王文倩，陈军，曹长群，王玥，王向东. 2019. 中国二叠纪综合地层和时间框架. 中国科学·地球科学, 49(1): 160-193。）

38.    Yuan, D.X., Shen, S.Z., Henderson, C.M., Chen, J., Zhang, H., Zheng, Q.F. & Wu, H.C. 2019. Integrative timescale for the Lopingian (Late Permian): A review and update from Shangsi, South China. Earth-Science Reviews 188: 190-209.

39.    Chen, J.T., Montañez, I.P., Qi, Y.P., Shen, S.Z. & Wang, X.D. 2018. Strontium and carbon isotopic evidence for decoupling of pCO2 from continental weathering at the apex of the late Paleozoic glaciation. Geology 46: 395-398.

40.    Yuan, D.X., Zhang, Y.C. & Shen, S.Z.* 2018. Conodont succession and reassessment of major events around the Permian-Triassic boundary at the Selong Xishan section, southern Tibet, China. Global and Planetary Change 161: 194-210.

41.    Wang, W.Q., Garbelli, C., Zheng, Q.F., Chen, J., Liu, X.C., Wang, W. & Shen, S.Z.* 2018. Permian ⁸⁷Sr/⁸⁶Sr chemostratigraphy from carbonate sequences in South China. Palaeogeography, Palaeoclimatology, Palaeoecology 500:84-94.

42.    Xu, H.P., Cao, C.Q., Yuan, D.X., Zhang, Y.C. & Shen, S.Z.* 2018. Lopingian (Late Permian) brachiopod faunas from the Qubuerga Formation at Tulong and Kujianla in the Mt. Everest area of southern Tibet, China. Rivista Italiana di Paleontologia e Stratigrafia 124: 139-162.

43.    Shen, S.Z. 2018. Global Permian brachiopod biostratigraphy: An overview. Special Publications of the Geological Society of London 450:289-320.

44.    Garbelli, C., Angiolini, L. & Shen, S.Z. 2017. Biomineralization and global change: A new perspective for understanding the end-Permian extinction. Geology 45:19-22.

45.    Godbold, A., Schoepfer, S., Shen, S.Z. & Henderson, C.M. 2017. Precarious ephemeral refugia during the earliest Triassic. Geology 45:607-610.

46.    Liu, S.A., Wu, H.C., Shen, S.Z., Jiang, G.Q., Zhang, S.H., Lv, Y.W., Zhang, H. & Li, S.G. 2017. Zinc isotope evidence for intensive magmatism immediately before the end-Permian mass extinction. Geology 45: 343-346.

47.    Yuan, D.X., Shen, S.Z. & Henderson, C.M. 2017. Revised Wuchiapingian conodont taxonomy and succession of South China. Journal of Paleontology 91:1199-1219.

48.    Wu, Q., Ramezani, J., Zhang, H., Wang, T.T., Yuan, D.X., Mu, L., Zhang, Y.C., Li, X.H. & Shen, S.Z. 2017. Calibrating the Guadalupian Series (Middle Permian) of South China. Palaeogeography, Palaeoclimatology, Palaeoecology 466: 361-372.

49.    沈树忠, 张华. 2017. 什么引起五次生物大灭绝？科学通报 62(11):1119-1135.

50.    Yuan, D.X., Zhang, Y.C., Shen, S.Z., Henderson, C.M., Zhang, Y.J., Zhu, T.X., An, X.Y. & Feng, H.Z. 2016. Early Permian conodonts from the Xainza area, central Lhasa Block, Tibet and their palaeobiogeographical and palaeoclimatic implications. Journal of Systematic Palaeontology 14:365-383.

51.    Chen, J., Shen, S.Z., Li, X.H., Xu, Y.G., Joachimski, M.M., Bowring, S.A., Erwin, D.H., Yuan, D.X., Chen, B., Zhang, H., Wang, Y., Cao, C.Q., Zheng, Q.F. & Mu, L. 2016. High-resolution SIMS oxygen isotope analysis on conodont apatite from South China and implications for the end-Permian mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 448: 26-38.

52.    Ke, Y., Shen, S.Z.*, Shi, G.R., Fan, J.X., Zhang, H., Qiao, L. & Zeng, Y. 2016. Global brachiopod palaeobiogeographical evolution from Changhsingian (Late Permian) to Rhaetian (Late Triassic). Palaeogeography, Palaeoclimatology, Palaeoecology 448: 4-25.

53.    Zhang, H., Cao, C.Q., Liu, X.L., Mu, L., Zheng, Q.F., Liu, F., Xiang, L., Liu, L.J. & Shen, S.Z.*. 2016. The terrestrial end-Permian mass extinction in South China. Palaeogeography, Palaeoclimatology, Palaeoecology 448: 108-124.

54.    Qiao Li & Shen, S.Z. 2015. A global review of the Late Mississippian (Carboniferous) Gigantoproductus (Brachiopoda) faunas and their paleogeographical, paleoecological, and paleoclimatic implications: Palaeogeography, Palaeoclimatology, Palaeoecology, 420:128-137.

55.    Shen, S.Z. & Bowring, S.A. 2014. The end-Permian mass extinction: A still unexplained catastrophe. National Science Review 1(4):492-495.

56.    Qiao, L. & Shen, S.Z. 2014. Global palaeobiogeography of brachiopods during the early Carboniferous—response to the global tectonic reconfiguration, ocean circulation, and climate changes. Gondwana Research 26:1173-1185.

57.    Yuan D.X., Shen, S.Z., Henderson, C.M., Chen, J. & Zhang, H. 2014. Revised conodont-based integrated high-resolution timescale for the Changhsingian Stage and end-Permian extinction interval at the Meishan sections, South China. Lithos 204: 220-245.

58.    Burgess, S.D., Bowring, S.A., Shen, S.Z. 2014. A new high-precision timeline for Earth’s most severe extinction. PNAS 111:3316-3321. 

59.    Wang, Y., Sadler, P.M., Shen, S.Z., Erwin, D.H., Zhang, Y.C., Wang, X.D., Wang, W., Crowley, J.L., Henderson, C.M. 2014. Quantifying the process and abruptness of the end-Permian mass extinction. Paleobiology 40:113-129 

60.    Shen, S.Z., Cao, C.Q., Zhang, H., Bowring, S.A., Henderson, C.M., Payne, J.L., Davydov, V.I., Chen, B., Yuan, D.X., Zhang, Y.C., Wang, W., Zheng, Q.F. 2013. High-resolution d¹³C_carb chemostratigraphy from latest Guadalupian through earliest Triassic in South China and Iran. Earth and Planetary Science letters, 375:156-165. 

61.    Shen, S.Z., Zhang, H., Shi, G.R., Li, W.Z., Xie, J.F., Mu, L., Fan, J.X. 2013. Early Permian (Cisuralian) global brachiopod palaeobiogeography. Gondwana Research 24:104-124. 

62.    Chen, B., Joachimski, M.M., Shen, S.Z., Lambert, L.L., Lai, X.L., Wang, X.D., Chen, J. & Yuan, D.X. 2013. Permian ice volume and palaeoclimate history: oxygen isotope proxies revisited. Gondwana Research 24: 77-89. 

63.    Zhang, Y.C., Shi, G.R., Shen, S.Z. 2013. A review of Permian stratigraphy, palaeobiogeography and palaeogeography of the Qinghai-Tibet Plateau. Gondwana Research 24:55-76. 

64.    Shen, S.Z., Yuan, D.X., Henderson, C.M., Tazawa, J. 2012. Implication of Kungurian (Early Permian) conodonts from Hatahoko, Japan, for correlation between the Tethyan and international timescales. Micropaleontology 58:505-522. 

65.    Shen, S.Z. 2012. Meishan sections in South China: The witness of the largest biological mass extinction during the Phanerozoic. Journal of Geography (Chigaku Zasshi) 121:570-678.

66.    Hinojosa, J.L., Brown, S.T., ChenJun, DePaolo, D.J., Paytan, A., Shen, S.Z. & Payne, J.L. 2012. Evidence for end-Permian ocean acidification from calcium isotopes in biogenic apatite. Geology 48:743-746.

67.    Joachimski, M.M., Lai, X.L., Shen, S.Z., Jiang, H.S., Luo, G.M., Chen, B., Chen, J., Sun Y.D. 2012. Climate warming in the latest Permian and the Permian–Triassic mass extinction. Geology 40:195-198.

68.    Zeng, J., Cao, C.Q., Davydov, V.I. & Shen, S.Z.* 2012. Carbon isotope chemostratigraphy and implications of palaeoclimatic changes during the Cisuralian (Early Permian) in the southern Urals, Russia. Gondwana Research 21:601-610.

69.    Shen, S.Z., Crowley, J.L., Wang, Y., Bowring, S.A., Erwin, D.H., Sadler, P.M., Cao, C.Q., Rothman, D.H., Henderson, C.M., Ramezani, J., Zhang, H., Shen, Y., Wang, X.D., Wang, W., Mu, L., Li, W.Z., Tang, Y.G., Liu, X.L., Liu, L.J., Zeng, Y., Jiang, Y.F., and Jin, Y.G. 2011. Calibrating the End-Permian Mass Extinction. Science 334:1367-1372.

70.    Shen, S.Z., Tazawa, J. & Miyake, Y. 2011. A Kungurian (Early Permian) Panthalassan brachiopod fauna from Hatahoko in the Mino belt, central Japan. Journal of Paleontology 85: 553-566.

71.    Shen, S.Z., Henderson, C.M., Bowring, S.A., Cao, C.Q., Wang, W., Wang, Y., Zhang, H., Zhang, Y.C. & Mu, L. 2010. High-resolution Lopingian (Late Permian) timescale of South China. Geological Journal 45:122-134.

72.    Shen, S.Z., Zhu, M.Y., Wang, X.D., Li, G.X., Cao, C.Q. & Zhang, H. 2010. A comparison of the biological, geological events and environmental backgrounds between the Neoproterozoic-Cambrian and Permian-Triassic transitions. Science in China Series D: Earth Sciences 53:1873-1884. (沈树忠、朱茂炎、王向东、李国祥、曹长群、张华, 2010.新元古代-寒武纪与二叠纪-三叠纪转折时期生物和地质事件及其环境背景之比较.中国科学D辑:地球科学, 40: 1228-1240).

73.    Shen, S.Z. & Clapham, M.E. 2009. Wuchiapingian (Lopingian, Late Permian) brachiopods from the Episkopi Formation at Hydra, Greece. Palaeontology 52:713-743.

74.    Shen, S.Z., Xie, J.F., Zhang, H. & Shi, G.R. 2009. Roadian-Wordian (Guadalupian, Middle Permian) global palaeobiogeography of brachiopods. Global and Planetary Change 65:166-181.

75.    Cao, C.Q., Love, G.D., Hays L.E., Wang, W., Shen, S.Z. & Summons, R.E. 2009. Biogeochemical evidence for euxinic oceans and ecological disturbance presaging the end-Permian mass extinction event. Earth and Planetary Science Letters, 281:188-201.

76.    Clapham, M.E., Shen, S.Z. & Bottjer, D.J. 2009. The double mass extinction revisited: Re-assessing the severity, selectivity, and causes of the end-Guadalupian biotic crisis (Late Permian). Paleobiology 35:33-51.

77.    Shen, S.Z. & Zhang, Y.C. 2008. Earliest Wuchiapingian (Lopingian, Late Permian) brachiopods in southern Hunan, south china: implications for the pre-Lopingian crisis and onset of Lopingian recovery/radiation. Journal of Paleontology 82: 924-937.

78.    Shen, S.Z., Cao, C.Q., Zhang, Y.C., Li, W.Z., Shi, G.R., Wang, Y., Wu, Y.S., Ueno, K., Henderson, C.M., Wang, X.D., Zhang, H., Wang, X.J. & Chen, J. 2010. End-Permian mass extinction and palaeoenvironmental changes across the Permian-Triassic boundary in the oceanic carbonate section in Neotethys. Global and Planetary Change, 73:3-14.

79.    Cao, C.Q., Wang, W., Liu, L.J., Shen, S.Z. & Summons, R.E. 2008. Two episodes of 13C-depletion in organic carbon in the latest Permian: Evidence from the terrestrial sequences in northern Xinjiang, China. Earth and Planetary Science Letters, 270:251-257.

80.    Shen, S.Z., Wang, Y., Henderson, C.M., Cao, C.Q. & Wang, W. 2007. Biostratigraphy and lithofacies of the Permian System in the Laibin-Heshan area of Guangxi, South China. Palaeoworld 16:120-139.

81.    Shen, S.Z., Zhang, H., Li, W.Z., Mu, L. & Xie, J.F. 2006. Brachiopod diversity pattern from Carboniferous to Triassic in South China. Geological Journal 41: 345-361.

82.    Shen, S.Z., Zhang, H., Shang, Q.H. & Li, W.Z. 2006. Permian lithostratigraphy and biostratigraphy of Northeast China. Journal of Asian Earth Sciences. 26:304-326.

83.    Shen, S.Z., Cao, C.Q., Henderson, C.M., Wang, X.D., Shi, G.R. Wang, W. & Wang Y. 2006. End-Permian mass extinction pattern in the northern peri-Gondwanan region. Palaeoworld 15:3-30.

84.    Jin, Y.G., Shen, S.Z.*, Henderson, C.M., Wang, X.D., Wang, W., Wang, Y., Cao, C.Q. & Shang, Q.H. 2006. The Global Stratotype Section and Point (GSSP) for the boundary between the Capitanian and Wuchiapingian stage (Permian). Episodes 29(4): 253-262.

85.    Jin Yugan, Wang Yue, C.M. Henderson, Shen, S.Z., Cao Changqun, Wang Wei, 2006. The Global Stratotype Section and Point (GSSP) for the base-Changhsingian Stage (Upper Permian). Episodes 29(3): 175-182.

86.    Shen, S.Z., Zhang, H., Li, W.Z., Mu, L. & Xie, J.F. 2006. Brachiopod diversity pattern from Carboniferous to Triassic in South China. Geological Journal 41(3): 345-361.

87.    Shen, S.Z., Grunt, T.G. & Jin, Y.G. 2004. A comparative study of Comelicaniidae Merla, 1930 (Brachiopoda, Athyridida) from the Lopingian (Late Permian) of South China and Transcaucasia in Azerbaijan and Iran. Journal of Paleontology 78:884-899.

88.    Shen, S.Z. & Shi, G.R. 2004. Capitanian (Late Guadalupian, Permian) global brachiopod palaeobiogeography and latitudinal diversity pattern. Palaeogeography, Palaeoclimatology, Palaeoecology 208: 235-262.

89.    Shen, S.Z., Cao, C.Q., Shi, G.R., Wang, X.D. & Mei, S.L. 2003. Lopingian (Late Permian) stratigraphy, sedimentation and palaeobiogeography in southern Tibet. Newsletters on Stratigraphy 39(2/3):157-179.

90.    Shen, S.Z., Shi, G.R. & Archbold, N.W., 2003. A Wuchiapingian (Late Permian) brachiopod fauna from an exotic block in the Indus-Tsangpo Suture zone, southern Tibet and its palaeobiogeographical and tectonic implication. Palaeontology 46(2):225-256.

91.    Shen, S.Z., Sun, D.L. & Shi, G.R. 2003. A Capitanian (late Middle Permian) brachiopod fauna from an exotic limestone block at Xiukang in Lhaze County of Tibet. Journal of Asian Earth Sciences 21(10):1125-1137.

92.    Shen, S.Z. & Shi, G.R. 2002. Paleobiogeographical extinction patterns of Permian brachiopods in the Asian-western Pacific Region. Paleobiology 28(4):449-463.

93.    Shen, S.Z., Archbold, N.W., Shi, G.R. & Chen, Z.Q. 2001. Permian brachiopods from the Selong Xishan section, Xizang (Tibet), China. Part 2: Palaeobiogeographical and palaeoecological implications, Spiriferida, Athyridida and Terebratulida. Geobios 34(2):157-182.

94.    Shen, S.Z. & Archbold, N.W. 2001. Chonetoidea (Brachiopoda) from the Lopingian of South China. Alcheringa 25(3-4): 327-350.

95.    Shen, S.Z., Archbold, N.W. & Shi, G.R., 2001. A Lopingian (Late Permian) brachiopod fauna from the Qubuerga Formation at Shengmi in the Mount Qomolangma region of Southern Xizang (Tibet), China. Journal of Palaeontology 75(2):274-283.

96.    Shen, S.Z. & Shi, G.R. 2000. Wuchiapingian (early Lopingian, Permian) global brachiopod palaeobiogeography: a quantitative approach. Palaeogeography, Palaeoclimatology, Palaeoecology 162:299-318.

97.    Shen, S.Z., Archbold, N.W., Shi, G.R. & Chen, Z.Q. 2000. Permian brachiopods from the Selong Xishan section, Xizang (Tibet), China. Part 1: Stratigraphy, Strophomenida, Productida and Rhynchonellida. Geobios 33(6):725-752.

98.    Shen, S.Z., Shi, G.R. & Zhu, K.Y. 2000. Early Permian brachiopods of Gondwana affinity from the Dingjiazhai Formation, Baoshan Block, western Yunnan, China. Rivista Italiana di Paleontologia e Stratigrafia 106(3):263-282.

99.    Shen, S.Z., Archbold, N.W. & Shi, G.R. 2000. Changhsingian (Late Permian) brachiopod palaeobiogeography. Historical Biology 15:121-134.

100.Shen, S.Z., Tazawa, J.I. & Shi, G.R. 1999. Peltichia from Asia: Taxonomy, biostratigraphy and palaeobiogeography. Journal of Paleontology 73(1): 47-61.

101.Shen, S.Z. & Jin Yugan. 1999. Brachiopods from the Permian-Triassic boundary beds at the Selong Xishan section, Xizang (Tibet), China. Journal of Asian Earth Science 17(4): 547-559.

102.Shen, S.Z. & Shi, G.R. 1996. Diversity and extinction patterns of Permian Brachiopoda of South China. Historical Biology 12:93-110.

103.Shen, S.Z., He, X.L. & Shi, G.R. 1995. Biostratigraphy and correlation of several Permian-Triassic boundary sections in southwestern China. Journal of Southeast Asian Earth Science 12(1-2):19-30．

104.Shen, S.Z., Fan, B.H., Zhang, C. & Zhang, X.P. 1994. A new species of permianellids, taxonomic and paleoecological significance. Geobios 27(4):477-485.